Two 49ers defensive stars, Pearsall open training camp on PUP list

Two 49ers defensive standouts were among those placed on the physically-unable-to-perform list Tuesday upon reporting to training camp in Santa Clara. Linebacker Dre Greenlaw and safety Talanoa Hufanga, coming off significant injuries sustained last season, will continue physical therapy until they receive medical clearance to return to action. The 49ers placed rookie wide receiver Ricky Pearsall (hamstring) on the non-football injury list last week. He sustained the injury while working out, general manager John Lynch said. Defensive end Drake Jackson also was placed on PUP as he continues to rehab from knee surgery last season. Players who are placed on the PUP list at the beginning of camp can be removed at any time. If a player opens the regular season on PUP, he must sit out at least the first four games. Greenlaw suffered a torn left Achilles in the first half of the Super Bowl as he sprung forward to enter the playing field for a defensive series. He underwent surgery four days later. Greenlaw is not expected to be ready for the 49ers' season opener on Monday, Sept. 9, against the New York Jets. "When I feel like I can go out there and I really contribute and bring...